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1、.,忍冬科Caprifoliaceae,.,中国植物志:13(12)/500(200+),FOC:5(66) /5(207),APG:42/890,Largely temperate/warm temperate in the northern hemisphere, some montane tropics (exp. Valerianoideae), but neither the Antipodes nor the Pacific.,大洋洲,太平洋,.,五福花科,锦带花亚科,败酱亚科,川续断亚科,忍冬亚科,北极花亚科,刺续断亚科,六道木属,.,中国植物志,接骨木属 Sambucus 荚蒾

2、属 Viburnum 莛子藨属 Triosteum 七子花属 Heptacodium 毛核木属 Symphoricarpos 鬼吹箫属 Leycesteria 忍冬属 Lonicera 北极花属 Linnaea 蝟实属 Kolkwitzia 六道木属 Zabelia 糯米条属 Abelia 双盾木属 Dipelta 锦带花属 Weigela,FOC,接骨木属 Sambucus 荚蒾属 Viburnum 莛子藨属 Triosteum 七子花属 Heptacodium 毛核木属 Symphoricarpos 鬼吹箫属 Leycesteria 忍冬属 Lonicera 北极花属 Linnaea 蝟实

3、属 Kolkwitzia 六道木属 Zabelia 糯米条属 Abelia 双盾木属 Dipelta 双六道木属 Diabelia 锦带花属 Weigela,五福花科 Adoxaceae,北极花科Linnaeaceae,锦带花科 Diervillaceae,忍冬科 Caprifoliaceae,双六道木属 Diabelia,.,忍冬科Cymes with a pair of bracts and 2 pairs of bracteoles,occasionally accrescent in fruit, rarely absent. anthers free, 2-celled,Ovary

4、inferior,Fruit a berry, a drupe with 2-5 pyrenes, or a leathery achene. Seeds 1 to many;,五福花科Inflorescences terminal panicles or umbellate, spicate, or capitate cymes.anthers free, 2-celled,Ovary semi-inferior to inferior,Fruit drupes; seeds 1 or 3-5.,锦带花科:Ovary inferior, with a globose gland on one

5、 side of upper part, Fruit a 2-valved dehiscent capsule,北极花科:bracts leaflike or reduced and located at base of ovary and forming an epicalyx.,.,1. 花序由聚散合成伞形、伞房式或圆锥式;柱头常2-3裂;花冠不具蜜腺;花药内向或外向五福花科 1. 花序非上述情况,若为圆锥花序则花柱细长;柱头大多为头状,很少分裂;花冠具蜜腺;花药内向。 2. 蒴果锦带花科 2. 核果。 3. 聚伞花序有一对苞片和两对小苞片,稀缺;种子1到多数忍冬科 3. 苞片叶状,着生在

6、子房基部形成萼状总苞;种子1-颗北极花科,.,五福花科Adoxaceae,1. Leaves simple; ovary 1-loculed; drupes 1-seeded; shrubs or small treesViburnum 1. Leaves compound; ovary 3-5-loculed; drupes usually 3-5-seeded; shrubs or small treesSambucus,.,五福花科Adoxaceae-荚蒾属 Viburnum,单叶,核果1种子,花药内向,叶柄及花冠筒被星状毛,花芳香,V. dilatatum 荚蒾,.,V. macroc

7、ephalum 绣球荚蒾,V. odoratissimum 珊瑚树,叶革职,不育花,.,五福花科Adoxaceae-接骨木属 Sambucus,S. williamsii接骨木,奇数羽状复叶,花药外向,核果浆果状,.,Adoxa moschatellina 五福花,草本,复叶,花,雄蕊2轮,内轮退化,外轮分裂为2半蕊花药外向,.,中国植物志,接骨木属 Sambucus 荚蒾属 Viburnum 莛子藨属 Triosteum 七子花属 Heptacodium 毛核木属 Symphoricarpos 鬼吹箫属 Leycesteria 忍冬属 Lonicera 北极花属 Linnaea 蝟实属 Ko

8、lkwitzia 六道木属 Zabelia 糯米条属 Abelia 双盾木属 Dipelta 锦带花属 Weigela,FOC,接骨木属 Sambucus 荚蒾属 Viburnum 莛子藨属 Triosteum 七子花属 Heptacodium 毛核木属 Symphoricarpos 鬼吹箫属 Leycesteria 忍冬属 Lonicera 北极花属 Linnaea 蝟实属 Kolkwitzia 六道木属 Zabelia 糯米条属 Abelia 双盾木属 Dipelta 双六道木属 Diabelia 锦带花属 Weigela,五福花科 Adoxaceae,北极花科Linnaeaceae,锦带

9、花科 Diervillaceae,忍冬科 Caprifoliaceae,双六道木属 Diabelia,.,忍冬科 Caprifoliaceae,1. Flowers single and opposite in a spike 2. Ovaries 5, 7, or 8-locular, all locules with many ovules. Leycesteria鬼吹箫属 2. Ovaries 4-locular, 2 locules with 1 fertile ovule, other 2 locules with many sterile ovules. Symphoricarpo

10、s毛核木属 1. Flowers paired (rarely single and then pedunculate; rarely in threes and then bracteoles fused into a cupule) or in whorls of 6 3. Flowers paired (rarely single and then pedunculate; rarely in threes and then bracteoles fused into a cupule). Lonicera忍冬属 3. Flowers in whorls of 6 4. Flowers

11、6 in a single whorl, with 2 pairs of tight involucral bracts forming a capitulum; capitula paniculate; shrubs with triplinerved leaves.Heptacodium七子花属,小苞片连合成杯状而包被萼筒,两对紧密缩合的苞片组成头状的圆锥花序,.,忍冬科 Caprifoliaceae-毛核木属Symphoricarpos,S. sinensis毛核木,灌木,花单生于短小、钻形苞片的腋内,形成顶生穗状花序,果顶端有一小喙,蓝黑色,具白霜,.,忍冬科 Caprifoliace

12、ae,1. Flowers single and opposite in a spike 2. Ovaries 5, 7, or 8-locular, all locules with many ovules. Leycesteria鬼吹箫属 2. Ovaries 4-locular, 2 locules with 1 fertile ovule, other 2 locules with many sterile ovules. Symphoricarpos毛核木属 1. Flowers paired (rarely single and then pedunculate; rarely i

13、n threes and then bracteoles fused into a cupule) or in whorls of 6 3. Flowers paired (rarely single and then pedunculate; rarely in threes and then bracteoles fused into a cupule). Lonicera忍冬属 3. Flowers in whorls of 6 4. Flowers 6 in a single whorl, with 2 pairs of tight involucral bracts forming

14、a capitulum; capitula paniculate; shrubs with triplinerved leaves.Heptacodium七子花属,小苞片连合成杯状而包被萼筒,两对紧密缩合的苞片组成头状的圆锥花序,.,忍冬科 Caprifoliaceae-七子花属 Heptacodium,One species,H. Miconioides 七子花,叶三出脉,宿存花萼被刺刚毛,花七朵,花冠稍二唇,.,1. Flowers single and opposite in a spike 2. Ovaries 5, 7, or 8-locular, all locules with

15、many ovules. Leycesteria鬼吹箫属 2. Ovaries 4-locular, 2 locules with 1 fertile ovule, other 2 locules with many sterile ovules.Symphoricarpos毛核木属 1. Flowers paired (rarely single and then pedunculate; rarely in threes and then bracteoles fused into a cupule) or in whorls of 6 3. Flowers paired (rarely

16、single and then pedunculate; rarely in threes and then bracteoles fused into a cupule). Lonicera忍冬属 3. Flowers in whorls of 6 4. Flowers 6 in a single whorl, with 2 pairs of tight involucral bracts forming a capitulum; capitula paniculate; shrubs with triplinerved leaves.Heptacodium七子花属,忍冬科 Caprifol

17、iaceae,小苞片连合成杯状而包被萼筒,两对紧密缩合的苞片组成头状的圆锥花序,.,4. Flowers 6 in a single whorl, with 2 pairs of tight involucral bracts forming a capitulum; capitula paniculate; shrubs with triplinerved leaves.Heptacodium七子花属 4. Inflorescence different, if flowers in a capitulum then plant a climber with perfoliate invol

18、ucral bracts and bilabiate flowers in 1-4 whorls 5. Herbs; leaves pinnatifid, otherwise perfoliate Triosteum莛子藨属 5. Shrubs or climbers; leaves entire, not perfoliate (involucral bracts sometimes perfoliate). 6. Shrubs; flowers funnelform Leycesteria鬼吹箫属 6. Climbers; flowers bilabiate Lonicera忍冬属,抱茎的

19、总苞,二唇的,羽裂的,漏斗状的,.,忍冬科 Caprifoliaceae-鬼吹箫属 Leycesteria,示蜜腺囊,穗状花序每节对生,子房7-8室,每室胚珠多数,L. gracilis纤细鬼吹箫,.,忍冬科 Caprifoliaceae-鬼吹箫属 Leycesteria,L. formosa鬼吹箫,花每节6朵, 苞片叶状,每轮6枚 花冠漏斗状 果具宿存萼齿,.,忍冬科 Caprifoliaceae,1. Flowers single and opposite in a spike 2. Ovaries 5, 7, or 8-locular, all locules with many ovu

20、les. Leycesteria鬼吹箫属 2. Ovaries 4-locular, 2 locules with 1 fertile ovule, other 2 locules with many sterile ovules. Symphoricarpos毛核木属 1. Flowers paired (rarely single and then pedunculate; rarely in threes and then bracteoles fused into a cupule) or in whorls of 6 3. Flowers paired (rarely single

21、and then pedunculate; rarely in threes and then bracteoles fused into a cupule). Lonicera忍冬属 3. Flowers in whorls of 6 4. Flowers 6 in a single whorl, with 2 pairs of tight involucral bracts forming a capitulum; capitula paniculate; shrubs with triplinerved leaves.Heptacodium七子花属,小苞片连合成杯状而包被萼筒,两对紧密缩

22、合的苞片组成头状的圆锥花序,.,4. Flowers 6 in a single whorl, with 2 pairs of tight involucral bracts forming a capitulum; capitula paniculate; shrubs with triplinerved leaves.Heptacodium七子花属 4. Inflorescence different, if flowers in a capitulum then plant a climber with perfoliate involucral bracts and bilabiate

23、 flowers in 1-4 whorls 5. Herbs; leaves pinnatifid, otherwise perfoliate Triosteum莛子藨属 5. Shrubs or climbers; leaves entire, not perfoliate (involucral bracts sometimes perfoliate). 6. Shrubs; flowers funnelform Leycesteria鬼吹箫属 6. Climbers; flowers bilabiate Lonicera忍冬属,抱茎的总苞,二唇的,羽裂的,漏斗状的,.,L. japon

24、ica忍冬,忍冬科 Caprifoliaceae-忍冬属 Lonicera,花二唇形,苞片,.,忍冬科 Caprifoliaceae-忍冬属 Lonicera,L. tragophylla盘叶忍冬,花序下1-2对叶连合成盘状,3朵花组成聚伞花序密集成头状花序,.,4. Flowers 6 in a single whorl, with 2 pairs of tight involucral bracts forming a capitulum; capitula paniculate; shrubs with triplinerved leaves.Heptacodium七子花属 4. Inf

25、lorescence different, if flowers in a capitulum then plant a climber with perfoliate involucral bracts and bilabiate flowers in 1-4 whorls 5. Herbs; leaves pinnatifid, otherwise perfoliate Triosteum莛子藨属 5. Shrubs or climbers; leaves entire, not perfoliate (involucral bracts sometimes perfoliate). 6.

26、 Shrubs; flowers funnelform Leycesteria鬼吹箫属 6. Climbers; flowers bilabiate Lonicera忍冬属,抱茎的总苞,二唇的,羽裂的,漏斗状的,.,忍冬科 Caprifoliaceae-莛子藨属 Triosteum,T. pinnatifidum莛子藨,草本,叶羽状深裂,果具宿存萼齿,3多花组成的聚伞花序对生,.,中国植物志,接骨木属 Sambucus 荚蒾属 Viburnum 莛子藨属 Triosteum 七子花属 Heptacodium 毛核木属 Symphoricarpos 鬼吹箫属 Leycesteria 忍冬属 Lo

27、nicera 北极花属 Linnaea 蝟实属 Kolkwitzia 六道木属 Zabelia 糯米条属 Abelia 双盾木属 Dipelta 锦带花属 Weigela,FOC,接骨木属 Sambucus 荚蒾属 Viburnum 莛子藨属 Triosteum 七子花属 Heptacodium 毛核木属 Symphoricarpos 鬼吹箫属 Leycesteria 忍冬属 Lonicera 北极花属 Linnaea 蝟实属 Kolkwitzia 六道木属 Zabelia 糯米条属 Abelia 双盾木属 Dipelta 双六道木属 Diabelia 锦带花属 Weigela,五福花科 Ad

28、oxaceae,北极花科Linnaeaceae,锦带花科 Diervillaceae,忍冬科 Caprifoliaceae,双六道木属 Diabelia,.,北极花科Linnaeaceae,1.Petioles of opposite leaf pairs dilated and connate at base, enclosing axillary buds; corolla hypocrateriform, with a cylindrical tube and 4 or 5 spreading lobes; flowers in 3-flowered cymes or paired; s

29、tigmas green and mucilaginousZabelia六道木属 1.Petioles of opposite leaf pairs not dilated, axillary buds exposed; corolla funnelform or bilabiate with 5 spreading lobes; flowers paired or single; stigmas white and not mucilaginous. 2. Two bracts accrescent and becoming winglike in fruit; ovary 4-locula

30、r with 2 fertile loculesDipelta双盾木属 2. Bracts not winglike in fruit; ovary 3- or 4-locular with 1 or 2 fertile locules.,扩大,合生,高脚碟形,圆柱形的,分泌粘液,漏斗形,.,北极花科Linnaeaceae-六道木属 Zabelia,TheZ. bifloraspecies complex,花冠高脚碟形,柱头绿色,叶柄基部膨大合生,花被裂片4,宿存花萼疏被纤毛,.,TheZabelia bifloraspecies complex was revised by Nakai (J

31、. Jap. Bot. 13(8): 1-12. 1937) who distinguished 11 species. We currently recognize two species (Zabelia biflora(Turczaninow) Makino六道木 and Zabelia dielsii(Graebner) Makino南方六道木) in China. Most of the differences between taxa were based on the length of the peduncles, bracts, and bracteoles, which a

32、re highly variable. Classification ofZabelia biflorais complex and needs to be investigated.,.,北极花科Linnaeaceae-六道木属 Zabelia,Z. triflora醉鱼草状六道木,宿存萼片 密被毛,花被裂片5,.,北极花科Linnaeaceae,1.Petioles of opposite leaf pairs dilated and connate at base; corolla hypocrateriform, with a cylindrical tube and 4 or 5 s

33、preading lobes; flowers in 3-flowered cymes or paired; stigmas green and mucilaginousZabelia六道木属 1.Petioles of opposite leaf pairs not dilated; corolla funnelform or bilabiate with 5 spreading lobes; flowers paired or single; stigmas white and not mucilaginous. 2. Two bracts accrescent and becoming

34、winglike in fruit; ovary 4-locular with 2 fertile loculesDipelta双盾木属 2. Bracts not winglike in fruit; ovary 3- or 4-locular with 1 or 2 fertile locules.,扩大,合生,高脚碟形,圆柱形的,分泌粘液,漏斗形,.,北极花科Linnaeaceae-双盾木属 Dipelta,D. floribunda 双盾木,小苞片盾状,膜质,宿存而增大成翅状,花冠稍二唇形,.,北极花科Linnaeaceae,1.Petioles of opposite leaf pa

35、irs dilated and connate at base; corolla hypocrateriform, with a cylindrical tube and 4 or 5 spreading lobes; flowers in 3-flowered cymes or paired; stigmas green and mucilaginousZabelia六道木属 1.Petioles of opposite leaf pairs not dilated; corolla funnelform or bilabiate with 5 spreading lobes; flower

36、s paired or single; stigmas white and not mucilaginous. 2. Two bracts accrescent and becoming winglike in fruit; ovary 4-locular with 2 fertile loculesDipelta双盾木属 2. Bracts not winglike in fruit; ovary 3- or 4-locular with 1 or 2 fertile locules.,扩大,合生,高脚碟形,圆柱形的,分泌粘液,漏斗形,.,3. Evergreen, creeping sub

37、shrubs; two bracts shieldlike and covered with sticky glandular hairs; inflorescence reduced to 2 single flowers at apices of branchlets (those can look like paired flowers but have more bracts and internodes separating them)Linnaea北极花属 3. Erect shrubs; bracts and inflorescence not as above. 4. Brac

38、ts and bracteoles covered with bristles, accrescent and becoming spongy, woody and fused to ovaries in fruit.Kolkwitzia蝟实属 4. Bracts and bracteoles without bristles, small and non-accrescent in fruit. 5. Flowers terminal and paired (flowers opening simultaneously), sometimes 1-3(-8) due to supernume

39、rary flowers axillary to bracteoles (more often so on repeat-bloom long shoots), spring floweringDiabelia双六道木属 5. Flowers axillary, paniculate, single or paired (flowers opening consecutively), summer or autumn floweringAbelia糯米条属,.,北极花科Linnaeaceae-北极花属 Linnaea,L. borealis北极花,One species,苞片盾状,密被粘性腺毛

40、,果实下垂,匍匐亚灌木,.,3. Evergreen, creeping subshrubs; two bracts shieldlike and covered with sticky glandular hairs; inflorescence reduced to 2 single flowers at apices of branchlets (those can look like paired flowers but have more bracts and internodes separating them)Linnaea北极花属 3. Erect shrubs; bracts

41、 and inflorescence not as above. 4. Bracts and bracteoles covered with bristles, accrescent and becoming spongy, woody and fused to ovaries in fruit.Kolkwitzia蝟实属 4. Bracts and bracteoles without bristles, small and non-accrescent in fruit. 5. Flowers terminal and paired (flowers opening simultaneou

42、sly), sometimes 1-3(-8) due to supernumerary flowers axillary to bracteoles (more often so on repeat-bloom long shoots), spring floweringDiabelia双六道木属 5. Flowers axillary, paniculate, single or paired (flowers opening consecutively), summer or autumn floweringAbelia糯米条属,.,K. amabilis蝟实,北极花科Linnaeaceae-蝟实属 Kolkwitzia,One species,花冠5裂,不等大,苞片小苞片密被刺刚毛,宿存,萼齿宿存,2朵花组成的聚伞花序伞房状,.,3. Evergreen, creeping subshrubs; two bracts shieldlike and covered with sticky glandular hairs; inflorescence reduced to 2 single flowers at apices of branchle

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